Anatomical Substrates of
Somatic Sensation

"The 2 principal somatic"
The 2 principal somatic
sensory systems:
Dorsal column-medial lemniscal system
Anterolateral system

Dorsal Column-Medial Lemniscal System
•Mediates mechanical sensations
-touch, limb position sense, vibration sense
•Well established; clinical & experimental

Anterolateral System
•Mediates protective sensations
-Pain
-Temperature (cold & warmth)
-Itch
•Not as definitively established as is the DC-ML system for touch

Perspective:
Peripheral somatic sensory receptors are sensitive to different stimulus qualities
Mechanical
Thermal (warm, cold)
Noxious (mechanical, thermal, polymodal)
Different receptor classes provide input to
different somatic sensory pathways
Differential thalamic and cortical localization

Dorsal column-medial lemniscal system: Mechanical sensations

Anterolateral system:
Pain, Thermal, Itch

Mechanoreceptors
are encapsulated

Nociceptors, thermoreceptors, & itch receptors are
bare nerve endings

Dorsal root ganglion neuron

Key Characteristics of DRG Neurons
Peripheral receptive field
Spatial area within which stimulation
activates the sensory neuron

Receptive field

CNS Neurons have Receptive Fields

Key Characteristics of DRG Neurons
Peripheral receptive field
Spatial area within which stimulation
activates the sensory neuron

Mechano-
receptor
receptive
fields

Mechano-
receptor
receptive
fields

Nociceptors, thermoreceptors, & itch receptors are
bare nerve endings

Nociceptors
respond to
noxious
stimuli

Peripheral
nerve
cross
section

Fiber Histogram: Sensory axon innervating the skin

Fiber Histogram: Sensory axon innervating a muscle

SC

Dermatomes

Dermatomes
overlap

Dermatomes
overlap

Dermatomes
overlap

Spinal cords

Receptor projections to systems

LgTermPatterns

DC lesion

Lg&SmTermPatterns

Lg&SmTermPatterns

Anterolateral system X

Spinal Hemisection

Syringomyelia

Somatotopy of spinal paths

Anterolateral
system

Somatic Sensory Thalamus

Slide 39

Slide 40

Slide 41

Brodmann’s
cytoarchitectonic
areas

Somatic Sensory Cortical Areas
for Mechanosensations

Slide 44

1° Somatic Sensory Cortex Output Systems

Cortical Pain Representations

Slide 47

Summary
Early morphological specialization of DRG neurons sets stage for separate mechanosensory and pain/temp/itch systems